How the Brain Regulates the Body’s Immune Response
Post by Meredith McCarty
The takeaway
The brain can modulate immune responses to prepare the immune system for potential threats or even induce placebo-like effects. This research reveals the functional circuitry underlying the coordination of behavioral and immunological responses in the mouse brain.
What's the science?
The immune system is responsible for taking protective action to protect the body against perceived external threats. The insular cortex (IC) of the brain is essential for integrating sensory information with bodily states. Prior work has implicated interactions between the insular cortex and the immune system, but the underlying functional dynamics remain unknown.
This week in Nature Neuroscience, Kayyal and colleagues use behavioral and experimental tools in mouse models to explore the role of IC in regulating immune responses.
How did they do it?
To study the interactions between the body’s immune response and sensory information, the authors utilize a conditioned immune response (CIR) task design. In this design, experimental mice are exposed to a conditioned stimulus (a specific scent) that is paired with an unconditioned stimulus that activates or inhibits the immune system. They learn the association between the scent and the unconditioned stimulus, and the researchers can study what is going on in the brain when this CIR is learned and tested.
To study brain activity, the researchers utilize retrograde labeling techniques to identify neurons within IC that project from anterior to posterior regions (aIC-to-pIC), or posterior to anterior regions (pIC-to-aIC). They tagged these populations to quantify the degree of connectivity, and changes in connectivity across various task conditions.
To study changes in the synaptic properties of the neurons, they conducted electrophysiological recordings from tagged neurons to quantify changes in excitability and inhibitory currents.
To test whether aIC-to-pIC neurons are necessary and sufficient for aspects of the measured CIR, they injected modified receptors into projecting neurons to allow them to experimentally activate or inhibit specific populations of neurons and measure changes in immune and behavioral responses.
What did they find?
When studying the immune response and behavior after CIR, they found the experimental mice had a strong aversion to the conditioned stimulus and an elevated immune response. This suggests that CIR primes the immune system for potential infection after exposure to harmful bacteria.
When comparing the neurons that project from aIC-to-pIC and vice versa, they found an increased percentage activation of aIC-to-pIC neurons following CIR, suggesting that these insular projections play a critical role in retrieving memory associated with the conditioned scent stimulus. When quantifying the excitatory to inhibitory ratio of projection neurons, they found reduced excitability and an increased number of active aIC-to-pIC projecting neurons, highlighting a potential mechanism by which information is flexibly retrieved during the retrieval of CIR-related memories.
Through selectively activating and inhibiting the aIC-to-pIC and pIC-to-aIC projections, they found that inhibiting the aIC-to-pIC pathway reduced learned aversive behavior. Their findings suggest that aIC regions encode taste and its conditioned response (immune system threat) and that the aIC-to-pIC pathway is necessary for the successful retrieval of the CIR. Their results highlight the importance of the aIC-to-pIC projecting neurons in modulating the body’s immune response following exposure to a conditioned aversive sensory stimulus.
What's the impact?
This study is the first to suggest a novel role for specific regions of the insular cortex in retrieving immune-related information and flexibly tuning behavioral responses. It was previously unknown how the functional connectivity of the insular cortex relates to the body’s immune response and how the brain and body interact to promote immune function.